@import url('https://fonts.googleapis.com/css2?family=Inter:wght@400;500;600&family=JetBrains+Mono:wght@400&display=swap');

*, *::before, *::after { box-sizing: border-box; margin: 0; padding: 0; }

:root {
    --canvas: #f7f7f4;
    --canvas-soft: #fafaf7;
    --surface-card: #ffffff;
    --surface-strong: #e6e5e0;
    --primary: #f54e00;
    --primary-active: #d04200;
    --on-primary: #ffffff;
    --ink: #26251e;
    --body: #5a5852;
    --body-strong: #26251e;
    --muted: #807d72;
    --muted-soft: #a09c92;
    --hairline: #e6e5e0;
    --hairline-soft: #efeee8;
    --hairline-strong: #cfcdc4;
    --success: #1f8a65;
    --error: #cf2d56;
    --radius-xs: 4px;
    --radius-sm: 6px;
    --radius-md: 8px;
    --radius-lg: 12px;
    --radius-xl: 16px;
    --radius-pill: 9999px;
}

html { scroll-behavior: smooth; }

body {
    background: var(--canvas);
    color: var(--body);
    font-family: 'Inter', system-ui, 'Helvetica Neue', Helvetica, Arial, sans-serif;
    font-size: 16px;
    line-height: 1.5;
    -webkit-font-smoothing: antialiased;
}

a { color: var(--ink); text-decoration: none; }
a:hover { color: var(--primary); }
ul[role="list"] { list-style: none; }
img { display: block; max-width: 100%; height: auto; }

.container {
    max-width: 1200px;
    margin: 0 auto;
    padding: 0 24px;
}

/* NAV */
.site-nav {
    position: sticky;
    top: 0;
    z-index: 100;
    background: var(--canvas);
    border-bottom: 1px solid var(--hairline);
    height: 64px;
}
.nav-inner {
    max-width: 1200px;
    margin: 0 auto;
    padding: 0 24px;
    height: 100%;
    display: flex;
    align-items: center;
    gap: 32px;
}
.nav-logo {
    display: flex;
    align-items: center;
    gap: 10px;
    flex-shrink: 0;
    color: var(--ink);
    font-size: 14px;
    font-weight: 500;
    letter-spacing: 0;
}
.nav-logo-mark {
    background: var(--primary);
    color: var(--on-primary);
    width: 32px;
    height: 32px;
    border-radius: var(--radius-sm);
    display: flex;
    align-items: center;
    justify-content: center;
    font-size: 13px;
    font-weight: 600;
    flex-shrink: 0;
}
.nav-links {
    display: flex;
    align-items: center;
    gap: 4px;
    list-style: none;
    margin-left: auto;
}
.nav-links li { position: relative; }
.nav-links a {
    display: block;
    padding: 8px 12px;
    border-radius: var(--radius-md);
    color: var(--ink);
    font-size: 14px;
    font-weight: 500;
    line-height: 1.4;
    transition: background 0.15s;
}
.nav-links a:hover { background: var(--surface-strong); color: var(--ink); }
.nav-dropdown {
    display: none;
    position: absolute;
    top: calc(100% + 4px);
    left: 0;
    background: var(--surface-card);
    border: 1px solid var(--hairline);
    border-radius: var(--radius-lg);
    min-width: 220px;
    list-style: none;
    padding: 8px;
    z-index: 200;
}
.nav-has-dropdown:hover .nav-dropdown,
.nav-has-dropdown:focus-within .nav-dropdown { display: block; }
.nav-dropdown a { padding: 8px 12px; font-weight: 400; white-space: nowrap; }
.nav-toggle {
    display: none;
    flex-direction: column;
    gap: 5px;
    background: none;
    border: none;
    cursor: pointer;
    padding: 8px;
    margin-left: auto;
}
.nav-toggle span {
    display: block;
    width: 22px;
    height: 2px;
    background: var(--ink);
    border-radius: 2px;
    transition: all 0.2s;
}

/* HERO */
.hero {
    padding: 80px 0 64px;
    border-bottom: 1px solid var(--hairline);
}
.hero-inner {
    max-width: 1200px;
    margin: 0 auto;
    padding: 0 24px;
    display: grid;
    grid-template-columns: 1fr 1fr;
    gap: 64px;
    align-items: center;
}
.hero-label {
    display: inline-block;
    background: var(--surface-strong);
    color: var(--muted);
    font-size: 11px;
    font-weight: 600;
    letter-spacing: 0.88px;
    text-transform: uppercase;
    padding: 4px 10px;
    border-radius: var(--radius-pill);
    margin-bottom: 20px;
}
.hero h1 {
    font-size: 48px;
    font-weight: 400;
    line-height: 1.1;
    letter-spacing: -1.5px;
    color: var(--ink);
    margin-bottom: 20px;
}
.hero-desc {
    font-size: 16px;
    line-height: 1.6;
    color: var(--body);
    margin-bottom: 32px;
    max-width: 480px;
}
.hero-image {
    position: relative;
    border-radius: var(--radius-lg);
    overflow: hidden;
    border: 1px solid var(--hairline);
}
.hero-image img {
    width: 100%;
    height: 360px;
    object-fit: cover;
}
.hero-image-caption {
    position: absolute;
    bottom: 0;
    left: 0;
    right: 0;
    background: linear-gradient(transparent, rgba(38,37,30,0.75));
    color: #fff;
    font-size: 13px;
    padding: 24px 16px 12px;
}

/* SECTION */
.section {
    padding: 80px 0;
}
.section + .section { border-top: 1px solid var(--hairline-soft); }
.section-label {
    display: inline-block;
    background: var(--surface-strong);
    color: var(--muted);
    font-size: 11px;
    font-weight: 600;
    letter-spacing: 0.88px;
    text-transform: uppercase;
    padding: 4px 10px;
    border-radius: var(--radius-pill);
    margin-bottom: 16px;
}
.section-title {
    font-size: 36px;
    font-weight: 400;
    line-height: 1.2;
    letter-spacing: -0.72px;
    color: var(--ink);
    margin-bottom: 12px;
}
.section-sub {
    font-size: 16px;
    color: var(--body);
    max-width: 560px;
    margin-bottom: 48px;
}

/* ARTICLE CARDS */
.article-grid {
    display: grid;
    grid-template-columns: repeat(3, 1fr);
    gap: 20px;
}
.article-card {
    background: var(--surface-card);
    border: 1px solid var(--hairline);
    border-radius: var(--radius-lg);
    overflow: hidden;
    display: flex;
    flex-direction: column;
    transition: border-color 0.15s;
}
.article-card:hover { border-color: var(--hairline-strong); }
.article-card-image { height: 200px; overflow: hidden; }
.article-card-image img { width: 100%; height: 100%; object-fit: cover; transition: transform 0.3s; }
.article-card:hover .article-card-image img { transform: scale(1.03); }
.article-card-body { padding: 24px; flex: 1; display: flex; flex-direction: column; }
.article-card-meta {
    font-size: 11px;
    font-weight: 600;
    letter-spacing: 0.88px;
    text-transform: uppercase;
    color: var(--muted);
    margin-bottom: 10px;
}
.article-card h3 {
    font-size: 18px;
    font-weight: 600;
    line-height: 1.35;
    color: var(--ink);
    margin-bottom: 10px;
}
.article-card p {
    font-size: 14px;
    color: var(--body);
    line-height: 1.55;
    flex: 1;
    margin-bottom: 20px;
}
.article-card-link {
    font-size: 14px;
    font-weight: 500;
    color: var(--primary);
    display: inline-flex;
    align-items: center;
    gap: 4px;
}
.article-card-link:hover { color: var(--primary-active); }

/* INFO BLOCKS */
.info-grid {
    display: grid;
    grid-template-columns: repeat(3, 1fr);
    gap: 20px;
}
.info-card {
    background: var(--surface-card);
    border: 1px solid var(--hairline);
    border-radius: var(--radius-lg);
    padding: 24px;
}
.info-card-icon {
    width: 40px;
    height: 40px;
    background: var(--surface-strong);
    border-radius: var(--radius-md);
    display: flex;
    align-items: center;
    justify-content: center;
    margin-bottom: 16px;
    font-size: 18px;
}
.info-card h3 {
    font-size: 18px;
    font-weight: 600;
    color: var(--ink);
    margin-bottom: 8px;
}
.info-card p { font-size: 14px; color: var(--body); line-height: 1.55; }

/* ARTICLE PAGE */
.article-hero {
    padding: 64px 0 0;
}
.article-hero-inner {
    max-width: 1200px;
    margin: 0 auto;
    padding: 0 24px;
}
.article-meta-row {
    display: flex;
    align-items: center;
    gap: 12px;
    margin-bottom: 20px;
    flex-wrap: wrap;
}
.article-category {
    background: var(--surface-strong);
    color: var(--muted);
    font-size: 11px;
    font-weight: 600;
    letter-spacing: 0.88px;
    text-transform: uppercase;
    padding: 4px 10px;
    border-radius: var(--radius-pill);
}
.article-date { font-size: 13px; color: var(--muted); }
.article-hero h1 {
    font-size: 42px;
    font-weight: 400;
    letter-spacing: -1px;
    line-height: 1.15;
    color: var(--ink);
    margin-bottom: 20px;
    max-width: 800px;
}
.article-hero-img {
    width: 100%;
    max-height: 480px;
    object-fit: cover;
    border-radius: var(--radius-lg);
    border: 1px solid var(--hairline);
    margin-top: 32px;
}
.article-img-credit {
    font-size: 12px;
    color: var(--muted-soft);
    margin-top: 6px;
}

.article-layout {
    max-width: 1200px;
    margin: 0 auto;
    padding: 48px 24px 80px;
    display: grid;
    grid-template-columns: 1fr 320px;
    gap: 64px;
    align-items: start;
}
.article-body h2 {
    font-size: 26px;
    font-weight: 400;
    letter-spacing: -0.325px;
    color: var(--ink);
    margin: 40px 0 16px;
    line-height: 1.25;
}
.article-body h2:first-child { margin-top: 0; }
.article-body h3 {
    font-size: 20px;
    font-weight: 600;
    color: var(--ink);
    margin: 28px 0 12px;
}
.article-body p {
    font-size: 16px;
    line-height: 1.7;
    color: var(--body);
    margin-bottom: 20px;
}
.article-body ul, .article-body ol {
    margin: 0 0 20px 20px;
    color: var(--body);
    font-size: 16px;
    line-height: 1.7;
}
.article-body li { margin-bottom: 6px; }
.article-body a { color: var(--primary); }
.article-body a:hover { color: var(--primary-active); }
.article-body strong { color: var(--ink); font-weight: 600; }

.article-highlight {
    background: var(--surface-card);
    border: 1px solid var(--hairline);
    border-left: 3px solid var(--primary);
    border-radius: var(--radius-md);
    padding: 20px 24px;
    margin: 28px 0;
}
.article-highlight p { margin: 0; font-size: 15px; }

.article-inline-img {
    width: 100%;
    border-radius: var(--radius-lg);
    border: 1px solid var(--hairline);
    margin: 28px 0 8px;
}

.article-sidebar { position: sticky; top: 84px; }
.sidebar-card {
    background: var(--surface-card);
    border: 1px solid var(--hairline);
    border-radius: var(--radius-lg);
    padding: 24px;
    margin-bottom: 20px;
}
.sidebar-card-title {
    font-size: 11px;
    font-weight: 600;
    letter-spacing: 0.88px;
    text-transform: uppercase;
    color: var(--muted);
    margin-bottom: 16px;
}
.sidebar-card ul { list-style: none; }
.sidebar-card ul li { border-top: 1px solid var(--hairline-soft); }
.sidebar-card ul li:first-child { border-top: none; }
.sidebar-card ul li a {
    display: block;
    padding: 10px 0;
    font-size: 14px;
    color: var(--body);
    transition: color 0.15s;
}
.sidebar-card ul li a:hover { color: var(--primary); }
.toc-list { list-style: none; }
.toc-list li { border-top: 1px solid var(--hairline-soft); }
.toc-list li:first-child { border-top: none; }
.toc-list a {
    display: block;
    padding: 8px 0;
    font-size: 14px;
    color: var(--body);
}
.toc-list a:hover { color: var(--primary); }

/* CONTACT FORM */
.contact-section { padding: 80px 0; }
.contact-inner {
    max-width: 720px;
    margin: 0 auto;
    padding: 0 24px;
}
.contact-form-card {
    background: var(--surface-card);
    border: 1px solid var(--hairline);
    border-radius: var(--radius-lg);
    padding: 40px;
}
.form-row { margin-bottom: 20px; }
.form-row label {
    display: block;
    font-size: 14px;
    font-weight: 500;
    color: var(--ink);
    margin-bottom: 6px;
}
.form-row input,
.form-row textarea {
    width: 100%;
    background: var(--surface-card);
    color: var(--ink);
    border: 1px solid var(--hairline-strong);
    border-radius: var(--radius-md);
    padding: 12px 16px;
    height: 44px;
    font-size: 16px;
    font-family: inherit;
    transition: border-color 0.15s;
    outline: none;
}
.form-row textarea { height: auto; min-height: 120px; resize: vertical; }
.form-row input:focus,
.form-row textarea:focus { border-color: var(--primary); }
.form-row input.field-error,
.form-row textarea.field-error { border-color: var(--error); }
.field-error-msg { font-size: 13px; color: var(--error); margin-top: 4px; display: none; }
.form-submit { margin-top: 8px; }
.form-status { margin-top: 16px; font-size: 14px; display: none; padding: 12px 16px; border-radius: var(--radius-md); }
.form-status.success { background: #e6f4ef; color: var(--success); border: 1px solid #b8dfd2; display: block; }
.form-status.error { background: #fce8ec; color: var(--error); border: 1px solid #f2bcc8; display: block; }

/* BUTTONS */
.btn-primary {
    display: inline-block;
    background: var(--primary);
    color: var(--on-primary);
    font-size: 14px;
    font-weight: 500;
    line-height: 1;
    padding: 10px 18px;
    height: 40px;
    border-radius: var(--radius-md);
    border: none;
    cursor: pointer;
    transition: background 0.15s;
}
.btn-primary:hover { background: var(--primary-active); color: var(--on-primary); }
.btn-primary:disabled { opacity: 0.6; cursor: not-allowed; }
.btn-secondary {
    display: inline-block;
    background: var(--surface-card);
    color: var(--ink);
    font-size: 14px;
    font-weight: 500;
    line-height: 1;
    padding: 10px 18px;
    height: 40px;
    border-radius: var(--radius-md);
    border: 1px solid var(--hairline-strong);
    cursor: pointer;
    transition: background 0.15s;
}
.btn-secondary:hover { background: var(--surface-strong); }

/* PAGE CONTENT (about/privacy/terms) */
.page-hero {
    padding: 64px 0 48px;
    border-bottom: 1px solid var(--hairline);
}
.page-hero-inner {
    max-width: 800px;
    margin: 0 auto;
    padding: 0 24px;
}
.page-hero h1 {
    font-size: 42px;
    font-weight: 400;
    letter-spacing: -1px;
    color: var(--ink);
    margin-bottom: 12px;
}
.page-hero p { font-size: 16px; color: var(--body); }
.page-content {
    max-width: 800px;
    margin: 0 auto;
    padding: 48px 24px 80px;
}
.page-content h2 {
    font-size: 26px;
    font-weight: 400;
    letter-spacing: -0.325px;
    color: var(--ink);
    margin: 40px 0 16px;
    line-height: 1.25;
}
.page-content h2:first-child { margin-top: 0; }
.page-content p {
    font-size: 16px;
    line-height: 1.7;
    color: var(--body);
    margin-bottom: 16px;
}
.page-content ul {
    margin: 0 0 16px 20px;
    color: var(--body);
    font-size: 16px;
    line-height: 1.7;
}
.page-content li { margin-bottom: 6px; }
.page-content a { color: var(--primary); }
.page-updated {
    display: inline-block;
    background: var(--surface-strong);
    color: var(--muted);
    font-size: 11px;
    font-weight: 600;
    letter-spacing: 0.88px;
    text-transform: uppercase;
    padding: 4px 10px;
    border-radius: var(--radius-pill);
    margin-bottom: 32px;
}

/* COOKIE BANNER */
.cookie-banner {
    position: fixed;
    bottom: 0;
    left: 0;
    right: 0;
    background: var(--surface-card);
    border-top: 1px solid var(--hairline);
    z-index: 999;
    display: none;
    padding: 16px 24px;
}
.cookie-inner {
    max-width: 1200px;
    margin: 0 auto;
    display: flex;
    align-items: center;
    gap: 24px;
    flex-wrap: wrap;
}
.cookie-inner p { font-size: 14px; color: var(--body); flex: 1; min-width: 200px; }
.cookie-inner a { color: var(--primary); }
.cookie-actions { display: flex; gap: 8px; flex-shrink: 0; }

/* FOOTER */
.site-footer {
    background: var(--canvas);
    border-top: 1px solid var(--hairline);
    padding: 64px 0 0;
}
.footer-inner {
    max-width: 1200px;
    margin: 0 auto;
    padding: 0 24px;
    display: grid;
    grid-template-columns: 2fr 1fr 1fr 1fr;
    gap: 40px;
    padding-bottom: 48px;
}
.footer-logo-mark {
    display: inline-flex;
    align-items: center;
    justify-content: center;
    background: var(--primary);
    color: var(--on-primary);
    width: 36px;
    height: 36px;
    border-radius: var(--radius-sm);
    font-size: 13px;
    font-weight: 600;
    margin-bottom: 16px;
}
.footer-tagline { font-size: 14px; color: var(--muted); line-height: 1.6; }
.footer-col-title {
    font-size: 11px;
    font-weight: 600;
    letter-spacing: 0.88px;
    text-transform: uppercase;
    color: var(--muted);
    margin-bottom: 16px;
}
.footer-col ul li { margin-bottom: 4px; }
.footer-col ul li a { font-size: 14px; color: var(--body); }
.footer-col ul li a:hover { color: var(--primary); }
.footer-bottom {
    border-top: 1px solid var(--hairline-soft);
    padding: 20px 24px;
    max-width: 1200px;
    margin: 0 auto;
    display: flex;
    justify-content: space-between;
    flex-wrap: wrap;
    gap: 8px;
}
.footer-bottom p { font-size: 13px; color: var(--muted); }
.footer-bottom a { color: var(--muted); }
.footer-bottom a:hover { color: var(--primary); }

/* RELATED SPECIES TABLE */
.species-table {
    width: 100%;
    border-collapse: collapse;
    font-size: 14px;
    margin: 24px 0;
}
.species-table th {
    background: var(--surface-strong);
    color: var(--ink);
    font-weight: 600;
    padding: 10px 16px;
    text-align: left;
    border-bottom: 1px solid var(--hairline);
}
.species-table td {
    padding: 10px 16px;
    border-bottom: 1px solid var(--hairline-soft);
    color: var(--body);
    vertical-align: top;
}
.species-table tr:last-child td { border-bottom: none; }

/* RESPONSIVE */
@media (max-width: 1024px) {
    .hero-inner { grid-template-columns: 1fr; gap: 40px; }
    .hero h1 { font-size: 40px; }
    .article-grid { grid-template-columns: repeat(2, 1fr); }
    .info-grid { grid-template-columns: repeat(2, 1fr); }
    .article-layout { grid-template-columns: 1fr; }
    .article-sidebar { position: static; }
    .footer-inner { grid-template-columns: 1fr 1fr; }
}
@media (max-width: 768px) {
    .hero h1 { font-size: 32px; letter-spacing: -0.5px; }
    .section-title { font-size: 28px; }
    .nav-links { display: none; flex-direction: column; align-items: flex-start; position: absolute; top: 64px; left: 0; right: 0; background: var(--canvas); border-bottom: 1px solid var(--hairline); padding: 16px 24px; gap: 0; }
    .nav-links.open { display: flex; }
    .nav-dropdown { position: static; border: none; box-shadow: none; padding: 4px 0 4px 16px; }
    .nav-has-dropdown:hover .nav-dropdown { display: none; }
    .nav-has-dropdown.open .nav-dropdown { display: block; }
    .nav-toggle { display: flex; }
    .article-grid { grid-template-columns: 1fr; }
    .info-grid { grid-template-columns: 1fr; }
    .contact-form-card { padding: 24px; }
    .footer-inner { grid-template-columns: 1fr; }
    .footer-bottom { flex-direction: column; gap: 4px; }
    .article-hero h1 { font-size: 30px; }
}
@media (max-width: 640px) {
    .hero { padding: 48px 0 40px; }
    .section { padding: 48px 0; }
    .hero h1 { font-size: 28px; }
}